An Annotated List of the Grasshoppers (Orthoptera: Acrididae, Romaleidae) of the Southeastern United States by Hill, JoVonn G. and M. E. Dakin

Research Article

An Annotated List of the Grasshoppers (Orthoptera: Acrididae, Romaleidae) of the Southeastern United States [pdf]

Hill¹, JoVonn G. and M. E. Dakin

¹Mississippi Entomological Museum, Box 9775, Mississippi State University, MS 39762
Email: jgh4@entomolgoy.msstate.edu

Received: 23-II-2011 Accepted: 29-IV-2011

Abstract: An annotated list, based on literature records, of the grasshopper species currently known from the southeastern United States is presented.
Key Words: Grasshoppers, southeastern United States, faunal list, Melanoplus

Introduction

The southeastern United States, including the states of Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, North Carolina, South Carolina, and Tennessee, for the purposes of this list, is often noted for its biological diversity. The region’s varied geography, climate, and geologic history have all played a role in producing a wide range of unique habitats, such as Appalachian boreal forests and balds, blackland prairies, limestone glades and barrens, long leaf pine savannahs, numerous classifications of hardwood and pine forests, sand hills, dunes, and wetlands. Studies of the region’s grasshopper fauna go back at least 150 years, but to date no comprehensive list of the fauna has been produced.
Researchers with the Mississippi Entomological Museum are currently sampling grasshoppers from the major ecoregions of the study area to describe the fauna more accurately and also to present more accurate species distribution records. The first step in producing such a product is to identify which species are already known from the region. To accomplish this, the literature was searched for reliable species records from the states listed above. The results of these searches are produced below in the form of an annotated checklist.

Results

Two families commonly referred to as grasshoppers are included in this list; the Acrididae (short-horned grasshoppers) and the Romaleidae (lubber grasshoppers), which is represented in the Southeast by a single species Romalea microptera (Beauvois). Currently, 160 total species-group names are known from the Southeast, with Florida and Georgia being the most diverse states (86 taxa each). Louisiana has the lowest number (39) of published species records. The total number of taxa known from each state is listed in Table 1. The most diverse subfamily of the Acrididae in the region is the Melanoplinae with 120 known taxa, followed by the Oedipodinae with 20, the Gomphocerinae with 18, the Cyrtacanthacridinae with 7, the Leptysminae with 2, and the Acridinae with 1. The most diverse genus is Melanoplus with 86 taxa being known from the region at present.

Table 1. State and number of taxa recorded in the scientific literature from each Southeastern state.

State	Number of Taxa recorded
Alabama	71
Arkansas	55
Florida	86
Georgia	86
Louisiana	39
Mississippi	50
North Carolina	76
South Carolina	55
Tennessee	55

Below, the taxa that are currently known from the Southeast are listed taxonomically by subfamily and then alphabetically within each subfamily. Each species is annotated by the abbreviation of the state in which they have been documented and the citation of the first or most reliable reference to support that record. Taxonomy follows Eades et al. (2010).

Acrididae

Gomphocerinae

Achurum carinatum (Walker): AL (Morse 1904b), FL (Scudder 1878), GA (Morse 1904b), MS (Blatchley 1920), SC (Rehn and Hebard 1916).

Ageneotettix deorum Scudder: AR (USDA 1956).

Amblytropidia mysteca (Saussure): AL (Morse 1907b), AR (Morse 1907b), FL (Scudder 1878), GA (Morse 1904b), LA (Otte 1981), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Saussure 1861b).

Boopedon auriventris McNeil: AR (McNeil 1899d).

Chloealtis conspersa Harris: AR (McNeil 1899d), NC (Morse 1904b), SC (Otte 1981), TN (USDA 1956).

Chorthippus curtipennis (Harris): NC (Morse 1904b), TN (Marcovitch 1920).

Dichromorpha elegans (Morse): AL (Morse 1904b), AR (Otte 1981), FL (Morse 1904b), GA (Morse 1904b), LA (Morse 1907b), MS (Morse 1907b), NC (Morse 1904b), SC (Rehn and Hebard 1916).

Dichromorpha viridis (Scudder): AL (Scudder 1878), AR (McNeil 1899d), FL (Scudder 1878), GA (Morse 1904b), LA (USDA 1956), MS (Morse 1907b), NC (Morse 1904b), SC (Stål 1873), TN (Morse 1904b).

Eritettix obscurus (Scudder): FL (Scudder 1878).

Eritettix simplex (Scudder): AL (Morse 1907b), AR (McNeil 1899d), GA (Morse 1904b), NC (Morse 1904b), SC (Rehn and Hebard 1916), TN (Marcovitch 1920).

Mermiria bivittata (Serville): AL (Morse 1907b), AR (Otte 1981), FL (Morse 1904b), GA (Scudder 1899), MS (Morse 1907b), NC (USDA 1957), SC (Blatchley 1920), TN (Otte 1981).

Mermiria intertexta Scudder: FL (Hebard 1916), GA (Scudder 1899), NC (Rehn and Hebard 1916), SC (Rehn and Hebard 1916).

Mermiria picta (Walker): AL (Blatchley 1920), AR (Otte 1981), FL (Morse 1904b), GA (Scudder 1877), LA (Otte 1981), MS (Blatchley 1920), NC (Scudder 1899), SC (Rehn and Hebard 1916), TN (Marcovitch 1920).

Orphulella pelidna (Burmeister): AL (Morse 1904b), AR (McNeil 1899d), FL (Scudder 1878), GA (Morse 1904b), LA (USDA 1957), MS (Morse 1907b), NC (Morse 1904), SC (Morse 1904b), TN (Morse 1904b).

Orphulella speciosa (Scudder): AL (Morse 1907b), AR (McNeil 1899d), LA (Blatchley 1920), MS (USDA 1957), NC (Morse 1904b), TN (USDA 1957).

Pseudopomala brachyptera (Scudder): MS (Hill 2005).

Stethophyma celata Otte: SC (Otte 1981).

Syrbula admirabilis (Uhler): AL (Morse 1904b), AR (McNeil 1899d), FL (Morse 1904b), GA (Morse 1904b), LA (Morse 1907b), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Morse 1904b).

Acridinae

Metaleptea brevicornis (Johannson): AL (Morse 1904b), AR (McNeil 1899d), FL (Morse 1904b), GA (Morse 1904b), LA (Morse 1907b), MS (Otte 1981), NC (Morse 1904b), SC (Rehn and Hebard 1916), TN (Morse 1904).

Oedipodinae

Arphia granulata (Sassure): AL (Dakin and Hays 1970), FL (Saussure 1884), GA (Rehn and Hebard 1904), MS (Blatchley 1920), NC (Rehn and Hebard 1916), SC (Rehn and Hebard 1916).

Arphia simplex Scudder: AR (Otte 1984), LA (Otte 1984).

Arphia sulphurea (Fabricius): AL (Morse 1907b), AR (Morse 1907b), FL (Rehn and Hebard 1904), GA (Morse 1904b), LA (Otte 1984), MS (Morse 1907b), NC (Morse 1904b), SC (Rehn and Hebard 1916), TN (Marcovitch 1920).

Arphia xanthoptera (Burmeister): AL (Morse 1904b), AR (Morse 1907b), FL (Morse 1904b), GA (Morse 1904b), LA (Otte 1984), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Morse 1904b).

Chortophaga austrailor Rehn & Hebard: AL (Dakin and Hays 1970), FL (Rehn and Hebard 1910a), GA (Rehn and Hebard 1910a), LA (Otte 1984), MS (Otte 1984).

Chortophaga viridifasciata (DeGeer): AL (Morse 1904b), AR (Morse 1907b), GA (Morse 1907), LA (USDA 1956), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Morse 1904b).

Dissosteira carolina (Linnaeus): AL (Morse 1904b), AR (Morse 1907b), FL (Scudder 1878), GA (Morse 1904b), LA (USDA 1956), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Morse 1904b).

Encoptolophus soridus (Burmeister): NC (Morse 1904b), TN (Morse 1904b).

Hippiscus ocelote (Saussure): AL (Morse 1904b), AR (Morse 1907b), FL (Scudder 1878), GA (Morse 1904b), LA (USDA 1956), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Morse 1904b).

Pardalophora apiculatus (Harris): NC (Rehn and Hebard 1910b), TN (Marcovitch 1920).

Pardalophora phoenicoptera (Burmeister): AL (Morse 1907b), AR (Morse 1907b), FL (Hebard 1915), GA (Rehn and Hebard 1904), LA (USDA 1957), MS (Morse 1907b), NC (Brimley 1908), SC (Rehn and Hebard 1916), TN (USDA 1957).

Psinidia fenestralis (Serville): AL (Rehn 1919), FL (Scudder 1878), GA (Morse 1904b), MS (Morse 1907b), NC (Morse 1904b), SC (Rehn and Hebard 1916).

Spharagemon bolli Scudder: AL (Morse 1907b), AR (Morse 1907b), FL (Morse 1904b), GA (Morse 1904b), LA (USDA 1957), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Morse 1904b).

Spharagemon crepitans (Saussure):GA (Saussure 1884), FL (Davis 1915).

Spharagemon cristatum (Scudder): AL (Dakin and Hays 1970), AR (Otte 1984), FL (Rehn and Hebard 1910), GA (Rehn and Hebard 1904), LA (Otte 1984), NC (Morse 1904b), SC (USDA 1957).

Spharagemon equale (Say): AR (USDA 1957).

Spharagemon marmorata picta (Scudder): AL (Dakin and Hayes 1970), FL (Scudder 1877), GA (Rehn and Hebard 1910a), MS (Morse 1907b), NC (Morse 1904b), SC (Rehn and Hebard 1916).

Trachyrhachys kiowa (Thomas): AL (USDA 1957), AR (Morse 1907b), GA (Morse 1904b), MS (Blatchley 1920), NC (USDA 1957), TN (Morse 1904b).

Trimerotropis maritima (Harris): AL (Morse 1904b), AR (McNeil 1901), FL (Morse 1904b), GA (Morse 1904b), LA (Morse 1907b), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Morse 1904b).

Trimerotropis saxatalis McNeil: AL (Dakin and Hays 1970), AR (McNeil 1901), GA (Morse 1904b), NC (Rehn and Hebard 1916), TN (Marcovitch 1920).

Leptysminae

Leptysma marginicollis (Serville): AL (Rehn and Eades 1961), FL (Scudder 1878), GA (Morse 1904b), LA (Morse 1907b), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Marcovitch 1920).

Stenacris vitreipennis (Marschall): AL (Rehn and Eades 1961), FL (Scudder 1878), GA (Marschall 1836), LA (Rehn and Eades 1961), NC (Rehn and Hebard 1916).

Cyrtacanthacridinae

Schistocerca alutacea (Harris): AL (Morse 1907b), AR (Morse 1907b), FL (Morse 1904b), GA (Morse 1904b), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Hubbell 1960).

Schistocerca americana (Drury): AL (Morse 1904b), AR (Morse 1907b), FL (Morse 1904b), GA (Morse 1904b), LA (Morse 1907b), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Morse 1904b).

Schistocera ceratiola Hubbell and Walker: FL (Hubbell and Walker 1928).

Schistocerca damnifica (Saussure): AL (Blatchley 1920), FL (Rehn and Hebard 1912), GA (Rehn and Hebard 1904), MS (Hill 2007), NC (Morse 1904b), SC (Rehn and Hebard 1916), TN (Saussure 1861a).

Schistocerca lineata Scudder: AR (Morse 1907b), GA (Song 2004), MS (Song 2004), NC (Hubbell 1960), TN (Hubbell 1960).

Schistocerca obscura (Fabricius):AL (USDA 1957), AR (Morse 1907b), FL (Rehn and Hebard 1910), GA (Rehn and Hebard 1910a), LA (Morse 1907b), MS (Morse 1907b), NC (Rehn and Hebard 1916), SC (Rehn and Hebard 1916), TN (Marcovitch 1920).

Schistocerca rubiginosa (Harris): AL (Hubbell 1960), FL (Morse 1904b), GA (Morse 1904b), LA (Hubbell 1960), MS (Hubbell 1960), NC (Song 2004), SC (Morse 1904b), TN (Song 2004).

Melanoplinae

Aptenopedes aptera aptera Scudder: FL (Scudder 1878).

Aptenopedes aptera borealis Hebard: GA (Rehn and Hebard 1916).

Aptenopedes aptera coquinae Hebard: FL (Hebard 1936).

Aptenopedes aptera saturiba Hebard: FL (Hebard 1936).

Aptenopedes aptera simplex Hebard: FL (Hebard 1936).

Aptenopedes hubbelli Hebard: FL (Hebard 1936).

Aptenopedes nigropicta Hebard: FL (Hebard 1936).

Aptenopedes robusta Hebard: FL (Hebard 1936).

Aptenopedes sphenariodes apalachee Hebard: AL (Hebard 1936), FL (Rehn and Hebard 1904), GA (Morse 1904b).

Aptenopedes sphenarioides clara Rehn: FL (Rehn 1902).

Aptenopedes sphenarioides sphenarioides Scudder: FL (Scudder 1878).

Booneacris variegata (Scudder): NC (Morse 1904b), TN (Marcovitch 1920).

Campylacantha olivacea (Scudder): AL (USDA 1956), AR (McNeil 1899a), GA (Rehn and Hebard 1916), MS (USDA 1956), SC (USDA 1956).

Dendrotettix australis (Morse): AL (Morse 1906), NC (Friauf 1957), TN (Friauf 1957).

Dendrotettix quercus Riley: AR (McNeil 1899a), TN (Friauf 1957).

Dendrotettix zimmermanni (Saussure): NC (Friauf 1957), SC (Friauf 1957).

Eotettix hebardi Rehn: AL (Dakin and Hays 1970), GA (Rehn 1906).

Eotettix palustris Morse: FL (Morse 1904a).

Eotettix pusillus Morse: AL (Dakin and Hays 1970), FL (Rehn and Hebard 1916), GA (Morse 1904a), SC (Morse 1904a).

Eotettix signatus Scudder: FL (Scudder 1897).

Gymnoscirtetes morsei Hebard: AL (Dakin and Hays 1970), FL (Hebard 1918).

Gymnoscirtetes pusillus Scudder: FL (Scudder 1897), GA (Morse 1904b).

Hesperotettix floridensis Morse: FL (Morse 1901), GA (Morse 1904b).

Hesperotettix gemmicula Hebard: AL (Dakin and Hays 1970), FL (Hebard 1918).

Hesperotettix osceola Hebard: FL (Hebard 1918).

Hesperotettix viridis brevipennis (Thomas): AL (Morse 1907b), FL (Morse 1904a), LA (USDA 1956), GA (Morse 1904b), MS (Hill 2007), SC (USDA 1956).

Hesperotettix viridis pratensis Scudder: AR (McNeil 1899a), TN (Marcovitch 1920).

Melanoplus acidocercus Hebard: GA (Hebard 1919).

Melanoplus acrophilus Hebard: GA (Otte 2002), NC (Hebard 1935).

Melanoplus alabamae Hebard: AL (Hebard 1920).

Melanoplus adelogyrus Hubbell: FL (Hubbell 1932).

Melanoplus angustipennis (Dodge): AR, (McNeil 1899c), GA (Rehn and Hebard 1916), LA (USDA 1956), NC (USDA 1956), SC (USDA 1956), TN (USDA 1956).

Melanoplus apalachicolae Hubbell: FL (Hubbell 1932).

Melanoplus attenuatus Scudder: GA (Rehn and Hebard 1916), NC (Scudder 1897), SC (Morse 1904b).

Melanoplus australis Morse: GA (Morse 1904a), SC (Rehn and Hebard 1916).

Melanoplus bispinosus Scudder: AL (Dakin and Hays 1970), AR (Morse 1907b), FL (Capinera et al. 1997), GA (Hill 2009).

Melanoplus bivitttatus (Say): AL (Morse 1907b), AR (McNeil 1899b), GA (Morse 1904b), LA (USDA 1956), MS (Hill 2007), NC (Morse 1904b), SC (USDA 1956), TN (Marcovitch 1920).

Melanoplus cantralli Dakin: AL (Dakin 1966).

Melanoplus carnegiei Morse: AL (Dakin and Hays 1970), GA (Morse 1904a), NC (Rehn and Hebard 1910b), SC (Morse 1904a).

Melanoplus celatus Morse: GA (Rehn and Hebard 1916), NC (Hebard 1937), SC (Hebard 1937), TN (Hebard 1937).

Melanoplus cherokee Hebard: GA (Hebard 1935), NC (Hebard 1935), SC (Hebard 1935).

Melanoplus clypeatus (Scudder): GA (Scudder 1877).

Melanoplus confusus Scudder:NC (Rehn and Hebard 1916).

Melanoplus dakini Hilliard: LA (Hillard 2001).

Melanoplus davisi Hebard: FL (Hebard 1918).

Melanoplus deceptus Morse: GA (Morse 1904a), NC (Morse 1904a), SC (Otte 2002), TN (Marcovitch 1920).
Melanoplus decoratus Morse: AL (Dakin and Hays 1970), GA (Morse 1904b), NC (Morse 1904a), TN (Marcovitch 1920).

Melanoplus decorus Rehn and Hebard: NC (Rehn and Hebard 1910b).

Melanoplus delaware Hebard: AL (Dakin and Hays 1970), GA (Hebard 1935), NC (Rehn and Hebard 1916), SC (Hebard 1935).

Melanoplus devius Morse: GA (Rehn and Hebard 1916), NC (Morse 1904a), SC (Rehn and Hebard 1916).

Melanoplus differentialis differentialis (Thomas): AL (Roberts 1942), AR (McNeil 1899b), LA (Morse 1907b), MS (Blatchley 1920), NC (Roberts 1942), TN (Morse 1904b).

Melanoplus differentialis nigricans Cockerell: AR (Roberts 1942), MS (Lago et al. 1988).

Melanoplus divergens Morse: GA (Hebard 1937), NC (Morse 1904a).

Melanoplus eurycercus Hebard: NC (Hebard 1935).

Melanoplus fasciatus (Walker): AR (McNeil 1899c).

Melanoplus femurrubrum (DeGeer): AL (Morse 1904b), AR (McNeil 1899c), GA (Morse 1904b), LA (USDA 1956), MS (Dakin 1985), NC (Morse 1904b), SC (USDA 1956), TN (Morse 1904b).

Melanoplus femurrubrum x propinquus: AL (Dakin 1985), LA (Dakin 1985).

Melanoplus foedus Scudder: AR (Morse 1907b).

Melanoplus forcipatus Hubbell: FL (Hubbell 1932).

Melanoplus foxi Hebard: GA (Hebard 1923).

Melanoplus furcatus Scudder: FL (Scudder 1897), GA (Rehn and Hebard 1916).

Melanoplus gracilis (Bruner): AL (Hebard 1935), AR (McNeil 1899c), MS (Hebard 1935), TN (Marcovitch 1920).
Melanoplus gurneyi Strohecker: FL (Strohecker 1960).

Melanoplus hubbelli Hebard: AL (Dakin and Hays 1970), GA (Hebard 1935), NC (Otte 2002), SC (Hebard 1935), TN (Hebard 1935).

Melanoplus impudicus Scudder: AL (Morse 1907b), AR (McNeil 1899c), FL (Gurney 1941), GA (Morse 1904b), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Gurney 1941).

Melanoplus indicifer Hubbell: FL (Hubbell 1932).

Melanoplus ingrami Hill: TN (Hill 2010).

Melanoplus keeleri keeleri (Thomas): AL (Dakin and Hays 1970), AR (McNeil 1899c), FL (Thomas 1874), GA (Morse 1904b), MS (Hill 2007), NC (Morse 1904b), SC (Morse 1904b), TN (Morse 1904b).

Melanoplus keeleri luridus (Dodge): AR (Morse 1904¹).

Melanoplus longicornis Saussure: GA (Hebard 1935).

Melanoplus mirus Rehn and Hebard: NC (Rehn and Hebard 1916).

Melanoplus morsei Blatchley: TN (Eades 1959).

Melanoplus nanciae Deyrup: FL (Deyrup 1996).

Melanoplus nubilus Rehn and Hebard: NC (Rehn and Hebard 1916).

Melanoplus nigrescens (Scudder): AL (Dakin and Hays 1970), GA (Scudder 1877), NC (Rehn and Hebard 1916).

Melanoplus oklahomae Hebard: AR (Hilliard 2001).

Melanoplus ordwayae Deyrup: FL (Deyrup 1996).

Melanoplus pachycercus Hebard: NC (Otte 2002).

Melanoplus packardii Scudder: AR (McNeil 1899c).

Melanoplus pegasus Hebard: GA (Hebard 1919).

Melanoplus ponderosus ponderosus (Scudder):AR (Morse 1907b), MS (USDA 1957), LA (USDA 1957), TN (USDA 1957).

Melanoplus ponderosus viola Thomas: AR (McNeil 1899b).

Melanoplus primaestivus Dakin: AL (Dakin 1966).

Melanoplus propinquus Scudder: AL (Dakin and Hays 1970), FL (Scudder 1897), GA (Morse 1904b), LA (Dakin 1985), MS (Morse 1907b), NC (Rehn and Hebard 1916), SC (Morse 1904b).

Melanoplus puer puer (Scudder): FL (Scudder 1878).

Melanoplus puer seminole Hubbell: FL (Hubbell 1932).

Melanoplus puer peninsularis Hubbell: FL (Hubbell 1932).

Melanoplus punctulatus arboreus Scudder: AL (Dakin and Hays 1970), FL (Rehn and Hebard 1916), GA (Rehn and Hebard 1916), MS (Lago et al. 1988).

Melanoplus punctulatus punctulatus (Scudder): FL (Peck et al. 1992), GA (Rehn and Hebard 1916), NC (Brimley 1908²).

Melanoplus pygmaeus Davis: FL (Davis 1915).

Melanoplus quercicola Hebard: FL (Hebard 1918), GA (Hill 2009).

Melanoplus querneus Rehn and Hebard: AL (Dakin and Hays 1970), FL (Peck et al 1992), GA (Rehn and Hebard 1916).

Melanoplus rotundipennis (Scudder): FL (Scudder 1878), GA (Rehn and Hebard 1916).

Melanoplus rusticus obovatipennis (Blatchley): AR (McNeil 1899c), TN (Marcovitch 1920).

Melanoplus sanguinipes vulturnus Gurney and Brooks: AL (Morse 1904b), AR (McNeil 1899c), FL (Morse 1904b), GA (Morse 1904b), LA (Morse 1907b), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b), TN (Morse 1904b).

Melanoplus scapularis Rehn and Hebard: FL (Hubbell 1932), GA (Rehn and Hebard 1916).

Melanoplus scudderi scudderi (Uhler): AL (USDA 1957), AR (McNeil 1899c), FL (Davis 1915), GA (Morse 1904b), MS (USDA 1957), LA (USDA 1957), NC (Morse 1904b), SC (Rehn and Hebard 1916), TN (Marcovitch 1920).

Melanoplus serrulatus Hebard: NC (Hebard 1937).

Melanoplus similis Morse: AL (Dakin and Hays 1970), NC (Morse 1904a), TN (Hebard 1920).

Melanoplus stegocercus Rehn and Hebard: GA (Rehn and Hebard 1916).

Melanoplus strumosus Morse: AL (Morse 1907b), FL (Morse 1904a), GA (Morse 1907), NC (Rehn and Hebard 1916), SC (Morse 1904b).

Melanoplus sylvaticus McNeil: AR (McNeil 1899c).

Melanoplus sylvestris Morse:NC (Morse 1904a).

Melanoplus symmetricus Morse: FL (Morse 1904a).

Melanoplus tepidus Morse: AL (Dakin and Hays 1970), FL (Capinera et al. 2001), MS (Morse 1906).

Melanoplus tequestae Hubbell: FL (Hubbell 1932).

Melanoplus tumidicercus Hubbell: GA (Hubbell 1932).

Melanoplus tunicae Hebard: AL (Dakin and Hays 1970), MS (Hebard 1920), LA (Hebard 1920), TN (Hebard 1920).

Melanoplus tribuloides Morse:AL (Morse 1906).

Melanoplus tribulus Morse: AL (Dakin and Hays 1970), GA (Morse 1904b).

Melanoplus vulnus Eades: MS (Morse 1907³), TN (Eades 1959).

Melanoplus walshii Scudder: AL (Dakin and Hays 1970), GA (Morse 1904b), NC (Morse 1904b), TN (Marcovitch 1920).

Melanoplus withlacoocheensisi Squitier and Deyrup: FL (Squitier et al. 1998).

Paratylotropidia beutenmuelleri Morse: NC (Morse 1907a).

Paratylotropidia brunneri Scudder: AR (McNeil 1899a).

Paroxya atlantica Scudder: AL (Blatchley 1920), FL (Scudder 1877), GA (Morse 1904b), MS (Morse 1907b), NC (Morse 1904b), SC (Morse 1904b).

Paroxya clavuliger (Serville): FL (Thomas 1874), GA (Morse 1904b), LA (Morse 1907b), MS (Morse 1907b), NC (Morse 1904b), SC (Davis 1978).

Paroxya hooseri (Blatchley): AL (Dakin and Hays 1970), LA (Morse 1907b).

Paroxya paroxyoides (Scudder): FL (Scudder 1878).

Paroxya recta Scudder: GA (Scudder 1877), FL (Scudder 1878).

Romaleidae

Romaleinae

Romalea microptera (Beauvois): AL (Morse 1907b), AR (Rehn and Grant 1959), FL (Scudder 1878), GA (Rehn and Hebard 1904), LA (Rehn and Grant 1959), MS (Morse 1907b), NC (Blatchley 1920), SC (Beauvois 1817), TN (Marcovitch 1920).

¹Morse was unable to satisfactorily determine which subspecies he had in Arkansas.

²only recorded as punctulatus

³recorded as M. morsei.

Acknowledgements

I thank Richard Brown and two anonymous reviewers for their comments on this manuscript. This article is approved for publication as Journal Article No. J11976 of the Mississippi Agricultural and Forestry Experiment Station.

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